Carabidae
Martin Luff & Andrew Duff
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Family CARABIDAE
Index to subfamilies
OMOPHRONINAE
CARABINAE
CICINDELINAE
LORICERINAE
ELAPHRINAE
SCARITINAE
BROSCINAE
TRECHINAE
HARPALINAE
BRACHININAE
OMOPHRON Latreille, 1802
SCOLYTUS Fabricius, 1790 non Müller, O.F., 1764
limbatum (Fabricius, 1777)
Tribe CARABINI
CALOSOMA Weber, 1801
inquisitor (Linnaeus, 1758)
sycophanta (Linnaeus, 1758) 1
CARABUS Linnaeus, 1758
Subgenus LIMNOCARABUS Géhin, 1876
clathratus Linnaeus, 1761
Subgenus EUCARABUS Géhin, 1885
arvensis Herbst, 1784
ssp. silvaticus Dejean, 1826
anglicanus Motschulsky, 1865
Subgenus CARABUS sensu stricto
granulatus Linnaeus, 1758
ssp. granulatus Linnaeus, 1758
ssp. hibernicus Lindroth, 1956
Subgenus MORPHOCARABUS Géhin, 1885
monilis Fabricius, 1792
consitus sensu auctt. non Panzer, 1809
insularis Born, 1908
Subgenus ARCHICARABUS Seidlitz, 1887
nemoralis Müller, O.F., 1764
Subgenus AUTOCARABUS Seidlitz, 1887
auratus Linnaeus, 1761 2
cancellatus Illiger, 1798 3
granulatus sensu Stephens, 1827 non Linnaeus, 1758
Subgenus HEMICARABUS Géhin, 1885
nitens Linnaeus, 1758
Subgenus TOMOCARABUS Reitter, 1896
convexus Fabricius, 1775 4
Subgenus OREOCARABUS Géhin, 1885
glabratus Paykull, 1790
ssp. lapponicus Born, 1908
Subgenus MESOCARABUS Thomson, C.G., 1875
problematicus Herbst, 1786
ssp. gallicus Géhin, 1885
catenulatus sensu auctt. non Scopoli, 1763
Subgenus CHAETOCARABUS Thomson, C.G., 1875
intricatus Linnaeus, 1761 Blue Ground Beetle
Subgenus MEGODONTUS Solier, 1848
violaceus Linnaeus, 1758 Violet Ground Beetle
ssp. purpurascens Fabricius, 1787
exasperatus sensu auctt. non Duftschmid, 1812
var. asperipennis Lapouge, 1921
ssp. sollicitans Hartert, 1907
britannicus Born, 1908
Tribe CYCHRINI
CYCHRUS Fabricius, 1794
caraboides (Linnaeus, 1758)
ssp. rostratus (Linnaeus, 1761)
Tribe NEBRIINI
LEISTUS Frölich, 1799
Subgenus POGONOPHORUS Latreille, 1802
montanus Stephens, 1827
rufomarginatus (Duftschmid, 1812)
spinibarbis (Fabricius, 1775)
Subgenus LEISTOPHORUS Reitter, 1905
fulvibarbis Dejean, 1826
Subgenus LEISTUS sensu stricto
ferrugineus (Linnaeus, 1758)
rufescens sensu auctt. partim non Fabricius, 1775
terminatus (Hellwig in Panzer, 1793)
rufescens (Fabricius, 1775)
praeustus (Fabricius, 1792)
NEBRIA Latreille, 1802
Subgenus EURYNEBRIA Ganglbauer, 1891
complanata (Linnaeus, 1767)
Subgenus PARANEBRIA Jeannel, 1937
livida (Linnaeus, 1758)
sabulosa (Fabricius, 1787)
lateralis (Fabricius, 1792)
Subgenus NEBRIA sensu stricto
brevicollis (Fabricius, 1792)
salina Fairmaire & Laboulbčne, 1854
degenerata Schaufuss, 1862
iberica Oliviera, 1876
klinckowstroemi Mjöberg, 1915
Subgenus BOREONEBRIA Jeannel, 1937
nivalis (Paykull, 1790)
rufescens (Ström, 1768)
gyllenhali (Schönherr, 1806)
balbii Bonelli, 1810
PELOPHILA Dejean, 1821
borealis (Paykull, 1790)
Tribe NOTIOPHILINI
NOTIOPHILUS Duméril, 1806
aesthuans Motschulsky, 1864
aestuans auctt.
pusillus Waterhouse, G.R., 1833 non von Schreber, 1759
aquaticus (Linnaeus, 1758)
pusillus (von Schreber, 1759)
strigifrons (Baudi, 1864)
blacki Edwards, J., 1913
biguttatus (Fabricius, 1779)
germinyi Fauvel, 1863
hypocrita sensu auctt. non Putzeys, 1866
palustris (Duftschmid, 1812)
quadripunctatus Dejean, 1826
quadriguttatus Fowler, 1886 lapsus calami
rufipes Curtis, 1829
substriatus Waterhouse, G.R., 1833
1 Rare immigrant.
2 Introduction, established in N. Essex and Berkshire.
3 Rare introduction.
4 Rare introduction?; recorded from Winstanley Park (Lancashire), in 1836.
CICINDELA Linnaeus, 1758
Subgenus CICINDELA sensu stricto
campestris Linnaeus, 1758 Green Tiger Beetle
hybrida Linnaeus, 1758
maritima Latreille & Dejean, 1822 Dune Tiger Beetle
sylvatica Linnaeus, 1758 Wood Tiger Beetle
Subgenus CYLINDERA Westwood, 1831
germanica Linnaeus, 1758
LORICERA Latreille, 1802
pilicornis (Fabricius, 1775)
coerulescens sensu auctt. non Linnaeus, 1758
ELAPHRUS Fabricius, 1775
Subgenus ELAPHRUS sensu stricto
cupreus Duftschmid, 1812
lapponicus Gyllenhal, 1810
uliginosus Fabricius, 1792
Subgenus TRICHELAPHRUS Semenov-Tian-Shanskij, 1926
riparius (Linnaeus, 1758)
DIACHEILA Motschulsky, 1845 †
arctica (Gyllenhal, 1810) †
polita (Faldermann, 1835) †
BLETHISA Bonelli, 1810
HELOBIUM Leach, 1815
multipunctata (Linnaeus, 1758)
DYSCHIRIUS Bonelli, 1810
Subgenus DYSCHIRIUS sensu stricto
angustatus (Ahrens, 1830)
obscurus (Gyllenhal, 1827)
thoracicus (Rossi, 1790)
arenosus Stephens, 1827
Subgenus DYSCHIRIODES Jeannel, 1941
aeneus (Dejean, 1825)
extensus Putzeys, 1845 1 Extinct?
elongatulus Dawson, 1856
globosus (Herbst, 1784)
gibbus (Fabricius, 1792)
impunctipennis Dawson, 1854
luedersi Wagner, 1915
aeneus sensu auctt. non Dejean, 1825
tristis sensu auctt. non Stephens, 1827
unicolor sensu auctt. non Motschulsky, 1844
nigricornis (Motschulsky, 1844) †
nitidus (Dejean, 1825)
politus (Dejean, 1825)
salinus Schaum, 1843
CLIVINA Latreille, 1802
collaris (Herbst, 1784)
contracta (Fourcroy, 1785)
fossor (Linnaeus, 1758)
1 Resident native, last recorded from Deal, E. Kent in 1940.
Tribe BROSCINI
BROSCUS Panzer, 1813
cephalotes (Linnaeus, 1758)
MISCODERA Eschscholtz, 1830
arctica (Paykull, 1798)
Tribe TRECHINI
PERILEPTUS Schaum, 1860
BLEMUS sensu Laporte, 1840 non Dejean, 1821
areolatus (Creutzer, 1799)
AEPUS Samouelle, 1819
AEPOPSIS Jeannel, 1922
marinus (Ström, 1783)
fulvescens Samouelle, 1819
robinii (Laboulbčne, 1849)
TRECHUS Clairville, 1806
BLEMUS sensu Stephens, 1827 non Dejean, 1821
Subgenus EPAPHIUS Samouelle, 1819
rivularis (Gyllenhal, 1810)
secalis (Paykull, 1790)
Subgenus TRECHUS sensu stricto
fulvus Dejean, 1831
lapidosus Dawson, 1849
obtusus Erichson, 1837
quadristriatus (Schrank, 1781)
minutus (Fabricius, 1781)
rubens sensu Clairville, 1806 non Fabricius, 1792
rubens (Fabricius, 1792)
paludosus (Gyllenhal, 1810)
subnotatus Dejean, 1831 1
THALASSOPHILUS Wollaston, 1854
longicornis (Sturm, 1825)
LASIOTRECHUS Ganglbauer, 1892
discus (Fabricius, 1792)
TRECHOBLEMUS Ganglbauer, 1892
micros (Herbst, 1784)
Tribe BEMBIDIINI
ASAPHIDION Des Gozis, 1886
TACHYPUS sensu auctt. non Weber, 1801
curtum (Heyden, 1870) 2
flavipes sensu auctt. partim non Linnaeus, 1761
flavipes (Linnaeus, 1761)
pallipes (Duftschmid, 1812)
stierlini (Heyden, 1880) 3
flavipes sensu auctt. partim non Linnaeus, 1761
BEMBIDION Latreille, 1802
Subgenus PHILOCHTHUS Stephens, 1828
aeneum Germar, 1824
biguttatum (Fabricius, 1779)
guttula (Fabricius, 1792)
iricolor Bedel, 1879
riparium sensu Fowler, 1886 partim non Olivier, 1795
lunulatum (Fourcroy, 1785)
riparium (Olivier, 1795)
mannerheimii Sahlberg, C.R., 1827
haemorrhoum sensu auctt. non Stephens, 1828
unicolor Chaudoir, 1850
Subgenus PRINCIDIUM Motschulsky, 1864
punctulatum Drapiez, 1821
Subgenus TESTEDIUM Motschulsky, 1864
bipunctatum (Linnaeus, 1761)
Subgenus ACTEDIUM Motschulsky, 1864
pallidipenne (Illiger, 1802)
ruficolle (Illiger, 1801) non Panzer, 1796
Subgenus NEJA Motschulsky, 1864
nigricorne Gyllenhal, 1827
Subgenus METALLINA Motschulsky, 1850
lampros (Herbst, 1784)
celere (Fabricius, 1792)
properans (Stephens, 1828)
velox Erichson, 1837 non Linnaeus, 1761
coeruleotinctum Reitter, 1908
cyaneotinctum Sharp, 1913
caeruleipenne Saunders, 1936
Subgenus EUPETEDROMUS Netolitzky, 1911
dentellum (Thunberg, 1787)
flammulatum Clairville, 1806
Subgenus NOTAPHUS Dejean, 1821
obliquum Sturm, 1825
semipunctatum Donovan, 1806
adustum Schaum, 1860
varium (Olivier, 1795)
ustulatum sensu Sturm, 1825 non Linnaeus, 1758
Subgenus NOTAPHEMPHANES Netolitzky, 1920
ephippium (Marsham, 1802)
Subgenus PLATAPHUS Motschulsky, 1864
prasinum (Duftschmid, 1812)
Subgenus TRICHOPLATAPHUS Netolitzky, 1914
BLEPHAROPLATAPHUS Netolitzky, 1920
virens Gyllenhal, 1827
Subgenus BEMBIDIONETOLITZKYA Strand, E., 1929
DANIELA Netolitzky, 1910 non Koch, 1891
atrocaeruleum (Stephens, 1828)
coeruleum Serville, 1826 4
geniculatum Heer, 1837/8
redtenbacheri Daniel, K., 1902
tibiale (Duftschmid, 1812)
Subgenus PERYPHUS Dejean, 1821
andreae (Fabricius, 1787)
ssp. bualei Jacquelin du Val, 1852
anglicanum Sharp, 1869
bruxellense Wesmael, 1835
rupestre sensu auctt. non Linnaeus, 1767
decorum (Zenker in Panzer, 1800)
deletum Audinet-Serville, 1821
nitidulum (Marsham, 1802) non (Schrank, 1781)
dalmatinum var. latinum sensu MacKechnie-Jarvis, 1932 non Netolitzky, 1911
femoratum Sturm, 1825
fluviatile Dejean, 1831
lunatum (Duftschmid, 1812)
maritimum Stephens, 1835
concinnum sensu auctt. non Stephens, 1828
?dorsuarium Bedel, 1879
monticola Sturm, 1825
petrosum Gebler, 1833 †
saxatile Gyllenhal, 1827
vectense Fowler, 1886
stephensii Crotch, 1866
affine (Stephens, 1835) non Say, 1823
testaceum (Duftschmid, 1812)
tetracolum Say, 1823
ustulatum sensu auctt. non Linnaeus, 1758
littorale sensu auctt. non Olivier, 1791
Subgenus NEPHA Motschulsky, 1864
callosum Küster, 1847 5
laterale Dejean, 1831 non Samouelle, 1819
genei Küster, 1847
quadriguttatum sensu (Illiger, 1798) non Fabricius, 1775
tetragrammum auctt. non (Chaudoir, 1846)
ssp. illigeri Netolitzky, 1914
Subgenus SYNECHOSTICTUS Motschulsky, 1864
stomoides Dejean, 1831
atroviolaceum sensu auctt. non Dufour, 1820
Subgenus PSEUDOLIMNAEUM Kraatz, 1888
inustum Jacquelin du Val, 1857 6
Subgenus LYMNAEUM Stephens, 1828
nigropiceum (Marsham, 1802)
Subgenus SEMICAMPA Netolitzky, 1910
gilvipes Sturm, 1825
schuppelii Dejean, 1831
schueppeli auctt.
Subgenus DIPLOCAMPA Bedel, 1896
assimile Gyllenhal, 1810
clarkii Dawson, 1849
fumigatum (Duftschmid, 1812)
Subgenus EMPHANES Motschulsky, 1850
minimum (Fabricius, 1792)
normannum Dejean, 1831
Subgenus BEMBIDION sensu stricto
LOPHA Dejean, 1821 non Bolten, 1798
humerale Sturm, 1825
quadrimaculatum (Linnaeus, 1761)
quadriguttatum (Fabricius, 1775)
quadripustulatum Audinet-Serville, 1821
quadriguttatum sensu (Olivier, 1795) non Fabricius, 1775
Subgenus TREPANEDORIS Netolitzky, 1918
doris (Panzer, 1796)
Subgenus TREPANES Motschulsky, 1864
articulatum (Panzer, 1795)
octomaculatum (Goeze, 1777) 7
sturmii (Panzer, 1804)
Subgenus PHYLA Motschulsky, 1844
obtusum Audinet-Serville, 1821
OCYS Stephens, 1828
BEMBIDION sensu auctt. partim non Latreille, 1802
harpaloides (Audinet-Serville, 1821)
rufescens (Guérin-Méneville, 1823)
quinquestriatus (Gyllenhal, 1810)
CILLENUS Samouelle, 1819
BEMBIDION sensu auctt. partim non Latreille, 1802
lateralis Samouelle, 1819
BRACTEON Bedel, 1879
BEMBIDION sensu auctt. partim non Latreille, 1802
CHRYSOBRACTEON Netolitzky, 1914
argenteolum (Ahrens, 1812) 8 Extinct
litorale (Olivier, 1790)
littorale (Olivier, 1791)
paludosum (Panzer, 1794)
Tribe TACHYINI
TACHYS Dejean, 1821
bistriatus (Duftschmid, 1812)
edmondsi Moore, 1956 9
piceus Edmonds, 1934 non Dalla Torre, 1877
micros (Fischer von Waldheim, 1828)
gregarius Chaudoir, 1846
scutellaris Stephens, 1828
ELAPHROPUS Motschulsky, 1839
TACHYS sensu auctt. partim non Dejean, 1821
parvulus (Dejean, 1831)
quadrisignatus (Duftschmid, 1812) 10
walkerianus (Sharp, 1913)
POROTACHYS Netolitzky, 1914
TACHYS sensu auctt. partim non Dejean, 1821
bisulcatus (Nicolai, 1822) 11 Extinct
fockii (Hummel, 1822)
Tribe POGONINI
POGONUS Dejean, 1821
chalceus (Marsham, 1802)
littoralis (Duftschmid, 1812)
luridipennis (Germar, 1822)
Tribe PATROBINI
PATROBUS Dejean, 1821
assimilis Chaudoir, 1844
clavipes Thomson, C.G., 1859
atrorufus (Ström, 1768)
excavatus (Paykull, 1790)
rufipes sensu (Duftschmid, 1812) non Fabricius, 1792
septentrionis Dejean, 1828
1 Established introduction. Possibly also resident native.
2 Added by Speight, Martinez & Luff (1986).
3 Added by Focarile (1964); see also Speight, Martinez & Luff (1986).
4 Added by Telfer (2001a).
5 Rare immigrant?; 1 specimen found at Woking Common (Surrey), in 1851.
6 Added by Levey & Pavett (1999).
7 Rediscovered by Jones (1992).
8 Former resident native, extinct; E. shore of L. Neagh, N. Ireland. Also temporary resident; E. Kent.
9 Endemic?
10 Rare introduction, probably with ships' ballast; 1 19th-Century record from South Shields (Durham).
11 Temporary introduction, probably with ships' ballast, extinct; recorded from South Shields (Durham), in 1863.
Tribe PTEROSTICHINI
STOMIS Clairville, 1806
pumicatus (Panzer, 1795)
POECILUS Bonelli, 1810
PTEROSTICHUS sensu auctt. partim non Bonelli, 1810
FERONIA Latreille, 1817
cupreus (Linnaeus, 1758)
affinis (Sturm, 1824)
erythropus (Dejean, 1828)
dinniki (Lutschnik, 1912)
caesica Donisthorpe, 1931
kugelanni (Panzer, 1797)
dimidiatus (Olivier, 1795) non Rossi, 1790
lepidus (Leske, 1785)
versicolor (Sturm, 1824)
caerulescens sensu auctt. non Linnaeus, 1758
PTEROSTICHUS Bonelli, 1810
Subgenus PTEROSTICHUS sensu stricto
cristatus (Dufour, 1820)
ssp. parumpunctatus Germar, 1824
Subgenus STEROPUS Stephens, 1828
aethiops (Panzer, 1796)
madidus (Fabricius, 1775)
concinnus (Sturm, 1818)
Subgenus PEDIUS Motschulsky, 1850
longicollis (Duftschmid, 1812)
inaequalis (Marsham, 1802) non Panzer, 1795
Subgenus LYPEROSOMUS Motschulsky, 1850
aterrimus (Herbst, 1784)
Subgenus ADELOSIA Stephens, 1835
macer (Marsham, 1802)
picimanus (Duftschmid, 1812)
Subgenus PLATYSMA Bonelli, 1810
niger (Schaller, 1783)
scotus (Jeannel, 1942)
Subgenus BOTHRIOPTERUS Chaudoir, 1838
adstrictus Eschscholtz, 1823
orinomum (Stephens, 1828)
vitreus (Dejean, 1828)
oblongopunctatus (Fabricius, 1787)
quadrifoveolatus Letzner, 1852
angustatus (Duftchmid, 1812) non Fabricius, 1787
Subgenus OMASEUS Stephens, 1828
melanarius (Illiger, 1798)
vulgaris sensu auctt. non Linnaeus, 1758
Subgenus PSEUDOMASEUS Chaudoir, 1838
anthracinus (Panzer, 1795)
gracilis (Dejean, 1828)
minor (Gyllenhal, 1827)
nigrita (Paykull, 1790)
rhaeticus Heer, 1837/8 1
nigrita sensu auctt. partim non Paykull, 1790
Subgenus LAGARUS Chaudoir, 1838
vernalis (Panzer, 1795)
crenatus auctt. non (Duftschmid, 1812)
Subgenus ARGUTOR Stephens, 1828
diligens (Sturm, 1824)
strenuus sensu Dawson, 1854 non Panzer, 1796
strenuus (Panzer, 1796)
erythropus (Marsham, 1802)
ABAX Bonelli, 1810
parallelepipedus (Piller & Mitterpacher, 1783)
ater (Villers, 1790)
striola (Fabricius, 1792)
parallelus (Duftschmid, 1812) 2 Extinct
Tribe PLATYNINI
SPHODRUS Clairville, 1806
leucophthalmus (Linnaeus, 1758)
planus (Fabricius, 1792)
CALATHUS Bonelli, 1810
Subgenus AMPHIGYNUS Haliday, 1841
rotundicollis Dejean, 1828
piceus sensu (Marsham, 1802) non Linnaeus, 1758
Subgenus CALATHUS sensu stricto
ambiguus (Paykull, 1790)
fuscus (Fabricius, 1792)
cinctus Motschulsky, 1850 3
mollis sensu auctt.partim non Marsham, 1802
erythroderus Gemminger & Harold, 1868
erratus (Sahlberg, C.R., 1827)
fulvipes sensu (Gyllenhal, 1810) non Fabricius, 1792
flavipes sensu (Duftschmid, 1812) non Fourcroy, 1785
fuscipes (Goeze, 1777)
cisteloides (Panzer, 1793)
melanocephalus (Linnaeus, 1758)
nubigena Haliday, 1838
micropterus (Duftschmid, 1812)
mollis (Marsham, 1802)
ochropterus sensu auctt. non Duftschmid, 1812
LAEMOSTENUS Bonelli, 1810
LAEMOSTHENES Agassiz, 1846
Subgenus LAEMOSTENUS sensu stricto
complanatus (Dejean, 1828) 4
Subgenus PRISTONYCHUS Dejean, 1828
terricola (Herbst, 1784)
subcyaneus (Illiger, 1801)
SYNUCHUS Gyllenhal, 1810
TAPHRIA Latreille, 1819
ODONTONYX Stephens, 1827
vivalis (Illiger, 1798)
nivalis (Panzer, 1797) non Paykull, 1790
PLATYDERUS Stephens, 1828
PLATYDERES Stephens, 1827 5
ruficollis (Marsham, 1802)
OLISTHOPUS Dejean, 1828
ODONTONYX sensu auctt. non Stephens, 1827
rotundatus (Paykull, 1790)
rotundicollis (Marsham, 1802)
SERICODA Kirby, 1837
AGONUM sensu auctt. partim non Bonelli, 1810
quadripunctata (De Geer, 1774) 6
ANCHOMENUS Bonelli, 1810
AGONUM sensu auctt. partim non Bonelli, 1810
dorsalis (Pontoppidan, 1763)
prasinus (Thunberg, 1784)
PARANCHUS Lindroth, 1974
AGONUM sensu auctt. partim non Bonelli, 1810
albipes (Fabricius, 1796)
ruficornis (Goeze, 1777) non De Geer, 1774
oblongus (Fabricius, 1792) non Fabricius, 1792
pallipes (Fabricius, 1801) non Fabricius, 1787
OXYPSELAPHUS Chaudoir, 1843
AGONUM sensu auctt. partim non Bonelli, 1810
ANCHUS LeConte, 1854
obscurus (Herbst, 1784)
?obscurus (Müller, O.F., 1776)
oblongus (Fabricius, 1792)
AGONUM Bonelli, 1810
Subgenus BATENUS Motschulsky, 1864
livens (Gyllenhal, 1810)
Subgenus EUROPHILUS Chaudoir, 1859
consimile (Gyllenhal, 1810) †
fuliginosum (Panzer, 1809)
gracile Sturm, 1824
micans Nicolai, 1822
piceum (Linnaeus, 1758)
scitulum Dejean, 1828
thoreyi Dejean, 1828
pelidnum sensu (Paykull, 1792) non Herbst, 1784
puellum Dejean, 1828
Subgenus AGONUM sensu stricto
afrum (Duftschmid, 1812)
moestum sensu auctt. non (Duftschmid, 1812)
emarginatum (Gyllenhal, 1827)
ericeti (Panzer, 1809)
gracilipes (Duftschmid, 1812)
elongatum Dejean, 1828
lugens (Duftschmid, 1812) 7
marginatum (Linnaeus, 1758)
muelleri (Herbst, 1784)
parumpunctatum (Fabricius, 1792)
chalybeum Sturm, 1824
nigrum Dejean, 1828
atratum sensu auctt. non Duftschmid, 1812
dahli (Preudhomme de Borre, 1879
sahlbergii (Chaudoir, 1850) 8 Extinct
archangelicum Sahlberg, J., 1874
sexpunctatum (Linnaeus, 1758)
versutum Sturm, 1824
viduum (Panzer, 1796)
moestum sensu Fowler & Donisthorpe, 1913 non Duftschmid, 1812
PLATYNUS Bonelli, 1810
AGONUM sensu auctt. partim non Bonelli, 1810
Subgenus LIMODROMUS Motschulsky, 1864
assimilis (Paykull, 1790)
?junceus (Scopoli, 1763)
angusticollis (Fabricius, 1801)
Tribe ZABRINI
ZABRUS Clairville, 1806
tenebrioides (Goeze, 1777)
gibbus (Fabricius, 1794)
AMARA Bonelli, 1810
Subgenus ZEZEA Csiki, 1929
TRIAENA LeConte, 1847 non Hübner, 1818
plebeja (Gyllenhal, 1810)
strenua Zimmermann, 1832
Subgenus AMARA sensu stricto
aenea (De Geer, 1774)
trivialis (Gyllenhal, 1810)
anthobia Villa & Villa, 1833
communis (Panzer, 1797)
vulgaris sensu Dawson, 1854 non Linnaeus, 1758
convexior Stephens, 1828
continua Thomson, C.G., 1873
curta Dejean, 1828
eurynota (Panzer, 1796)
acuminata (Paykull, 1798)
famelica Zimmermann, 1832
familiaris (Duftschmid, 1812)
lucida (Duftschmid, 1812)
lunicollis Schiödte, 1837
vulgaris sensu auctt. non Linnaeus, 1758
montivaga Sturm, 1825
nitida Sturm, 1825
ovata (Fabricius, 1792)
obsoleta sensu Dejean, 1828 non Duftschmid, 1812
adamantina Kolenati, 1845
similata (Gyllenhal, 1810)
obsoleta (Duftschmid, 1812)
spreta Dejean, 1831
tibialis (Paykull, 1798)
Subgenus CELIA Zimmermann, 1832
bifrons (Gyllenhal, 1810)
livida sensu Schiödte, 1841 non Fabricius, 1792
cursitans (Zimmermann, 1832) 9
fuscicornis (Zimmermann, 1832)
fusca Sturm in Dejean, 1828
complanata Dejean, 1828
infima (Duftschmid, 1812)
praetermissa (Sahlberg, C.R., 1827)
rufocincta (Sahlberg, C.R., 1827)
Subgenus PARACELIA Bedel, 1899
quenseli (Schönherr, 1806)
quenselii auctt.
Subgenus BRADYTUS Stephens, 1827
apricaria (Paykull, 1790)
consularis (Duftschmid, 1812)
fulva (Müller, O.F., 1776)
Subgenus PERCOSIA Zimmermann, 1832
equestris (Duftschmid, 1812)
patricia (Duftschmid, 1812)
CURTONOTUS Stephens, 1827
alpinus (Paykull, 1790)
aulicus (Panzer, 1796)
spinipes sensu Schiödte, 1841 non Linnaeus, 1758
convexiusculus (Marsham, 1802)
torridus (Panzer, 1797) †
Tribe HARPALINI
HARPALUS Latreille, 1802
Subgenus HAPLOHARPALUS Schauberger, 1926
froelichii Sturm, 1818
Subgenus HARPALUS sensu stricto
affinis (Schrank, 1781)
aeneus (Fabricius, 1775) non De Geer, 1774
anxius (Duftschmid, 1812)
attenuatus Stephens, 1828
consentaneus Dejean, 1829
cupreus Dejean, 1829 10 Extinct?
dimidiatus (Rossi, 1790)
caspius (von Steven, 1806)
honestus (Duftschmid, 1812) 11
ignavus (Duftschmid, 1812)
latus (Linnaeus, 1758)
erythrocephalus (Fabricius, 1787)
metallescens Rye, 1874
melancholicus Dejean, 1829
neglectus Audinet-Serville, 1821
pumilus Sturm, 1818
vernalis (Duftschmid, 1812) non Panzer, 1796
picipennis sensu auctt. non Duftschmid, 1812
funestus Audinet-Serville, 1821
quadripunctatus Dejean, 1829
seriepunctatus sensu Gyllenhal, 1827 non Sturm, 1818
montivagus Reitter, 1900
rubripes (Duftschmid, 1812)
sobrinus Dejean, 1829
rufipalpis Sturm, 1818
rufitarsis (Duftschmid, 1812) non Illiger, 1801
ignavus sensu auctt. Brit. non Duftschmid, 1812
serripes (Quensel in Schönherr, 1806)
servus (Duftschmid, 1812)
smaragdinus (Duftschmid, 1812)
discoideus Erichson, 1837
tardus (Panzer, 1796)
rufimanus (Marsham, 1802)
luteicornis sensu auctt. Brit. non Duftschmid, 1812
tenebrosus Dejean, 1829
ssp. centralis Schauberger, 1929
Subgenus PSEUDOOPHONUS Motschulsky, 1844
griseus (Panzer, 1797) 12
rufipes (De Geer, 1774)
ruficornis (Fabricius, 1775)
pubescens (Müller, O.F., 1776)
Subgenus PARDILEUS Des Gozis, 1882
calceatus (Duftschmid, 1812) 13
OPHONUS Dejean, 1821
HARPALUS sensu auctt. partim non Latreille, 1802
Subgenus OPHONUS sensu stricto
ardosiacus Lutschnik, 1922
rotundicollis Fairmaire & Laboulbčne 1854 non Kolenati, 1845
azureus (Fabricius, 1775)
subquadratus sensu auctt. Brit. non Kolenati, 1845
sabulicola (Panzer, 1796)
stictus (Stephens, 1828)
monticola Dejean, 1829
obscurus (Fabricius, 1792) non Müller, O.F., 1776
Subgenus METOPHONUS Bedel, 1895
cordatus (Duftschmid, 1812)
melletii Heer, 1837/8
rectangulus (Thomson, C.G., 1870)
championi Sharp, 1912
rupicoloides Sharp, 1912
brevicollis sensu Jeannel, 1942 ? Audinet-Serville, 1821
parallelus Dejean, 1829
melletii sensu Jeannel, 1942 non Heer, 1837/8
zigzag sensu auctt. Brit. non Costa, 1882
punctatulus (Duftschmid, 1812)
nitidulus sensu (Stephens, 1828) non Schrank, 1781
puncticeps (Stephens, 1828)
rectangulus sensu Sharp, 1912 non Thomson, C.G., 1870
angusticollis (Müller, J., 1921)
puncticollis (Paykull, 1798)
rufibarbis (Fabricius, 1792)
brevicollis sensu auctt. partim non Audinet-Serville, 1821
subpunctatus (Stephens, 1828)
seladon Schauberger, 1926
rupicola Sturm, 1818
schaubergerianus Puel, 1937
rufibarbis sensu auctt. non Fabricius, 1792
brevicollis sensu auctt. partim non Audinet-Serville, 1821
subsinuatus Rey, 1886 14
ANISODACTYLUS Dejean, 1829
binotatus (Fabricius, 1787)
spurcaticornis Dejean, 1829
nemorivagus (Duftschmid, 1812)
atricornis (Stephens, 1835)
poeciloides (Stephens, 1828)
pseudoaeneus sensu auctt. non Dejean, 1829
DIACHROMUS Erichson, 1837
germanus (Linnaeus, 1758) 15 Extinct
SCYBALICUS Schaum, 1862
oblongiusculus (Dejean, 1829) 16 Extinct?
STENOLOPHUS Dejean, 1821
AGONODERUS Dejean, 1829
comma (Fabricius, 1775) 17
plagiatus Gorham, 1901
mixtus (Herbst, 1784)
vespertinus (Panzer, 1796)
ziegleri (Panzer, 1809)
skrimshiranus Stephens, 1828
teutonus (Schrank, 1781)
vaporariorum sensu (Fabricius, 1775) non Linnaeus, 1758
anglicus Schiödte, 1861-63
BRADYCELLUS Erichson, 1837
caucasicus (Chaudoir, 1846)
collaris (Paykull, 1798) non Herbst, 1784
csikii Lacz˘, 1912
distinctus (Dejean, 1829)
harpalinus (Audinet-Serville, 1821)
ruficollis (Stephens, 1828)
similis (Dejean, 1829)
sharpi Joy, 1912
distinctus sensu Fowler, 1886 non Dejean, 1829
verbasci (Duftschmid, 1812)
DICHEIROTRICHUS Jacquelin du Val, 1855
gustavii Crotch, 1871
pubescens (Paykull, 1790) non Müller, O.F., 1776
obsoletus (Dejean, 1829)
TRICHOCELLUS Ganglbauer, 1892
cognatus (Gyllenhal, 1827)
placidus (Gyllenhal, 1827)
ACUPALPUS Latreille, 1829
brunnipes (Sturm, 1825)
dubius Schilsky, 1888
luridus sensu auctt. non Dejean, 1829
elegans (Dejean, 1829)
exiguus Dejean, 1829
flavicollis (Sturm, 1825)
luridus Dejean, 1829
meridianus (Linnaeus, 1761)
parvulus (Sturm, 1825)
dorsalis (Fabricius, 1787) non Pontoppidan, 1763
derelictus Dawson, 1854
ANTHRACUS Motschulsky, 1850
ACUPALPUS sensu auctt. partim non Latreille, 1829
consputus (Duftschmid, 1812)
Tribe LICININI
LICINUS Latreille, 1802
depressus (Paykull, 1790)
punctatulus (Fabricius, 1792)
silphoides sensu (Fabricius, 1792) non Rossi, 1790
BADISTER Clairville, 1806
Subgenus BADISTER sensu stricto
bullatus (Schrank, 1798)
bipustulatus (Fabricius, 1792) non Fabricius, 1775
meridionalis Puel, 1925
kineli Makolski, 1952
sodalis (Duftschmid, 1812)
humeralis Bonelli, 1813
unipustulatus Bonelli, 1813
Subgenus BAUDIA Ragusa, 1884
collaris (Motschulsky, 1860)
anomalus (Perris, 1866)
striatulus Hansen, 1944
dilatatus Chaudoir, 1837
peltatus (Panzer, 1797)
Tribe OODINI
OODES Bonelli, 1810
gracilis Villa, 1833 †
helopioides (Fabricius, 1792)
Tribe PANAGAEINI
PANAGAEUS Latreille, 1802
bipustulatus (Fabricius, 1775)
quadripustulatus Sturm, 1815
cruxmajor (Linnaeus, 1758)
Tribe CALLISTINI
CHLAENIUS Bonelli, 1810
CHLAENIELLUS Reitter, 1908
costulatus Motschulsky, 1859 †
nigricornis (Fabricius, 1787)
melanocornis Dejean, 1826
nitidulus (Schrank, 1781) 18
schrankii (Duftschmid, 1812)
sulcicollis (Paykull, 1798) †
tristis (Schaller, 1783)
holosericeus (Fabricius, 1787)
vestitus (Paykull, 1790)
CALLISTUS Bonelli, 1810
lunatus (Fabricius, 1775)
Tribe PERIGONINI
PERIGONA Laporte, 1835
nigriceps (Dejean, 1831) 19
Tribe ODACANTHINI
ODACANTHA Paykull, 1798
COLLIURIS sensu auctt. non De Geer, 1774
melanura (Linnaeus, 1767)
Tribe MASOREINI
MASOREUS Dejean, 1821
wetterhallii (Gyllenhal, 1813)
laticollis (Sturm, 1825)
Tribe LEBIINI
LEBIA Latreille, 1802
Subgenus LAMPRIAS Bonelli, 1810
chlorocephala (Hoffmann, J., 1803)
chrysocephala Motschulsky, 1864
cyanocephala (Linnaeus, 1758)
Subgenus LEBIA sensu stricto
cruxminor (Linnaeus, 1758)
marginata (Fourcroy, 1785) 20 Extinct
haemorrhoidalis (Fabricius, 1787)
scapularis (Fourcroy, 1785) 21 Extinct
turcica (Fabricius, 1787)
Tribe DEMETRIINI
DEMETRIAS Bonelli, 1810
RISOPHILUS Leach, 1815
Subgenus AETOPHORUS Schmidt-Göbel, 1846
imperialis (Germar, 1824)
Subgenus DEMETRIAS sensu stricto
atricapillus (Linnaeus, 1758)
monostigma Samouelle, 1819
unipunctatus (Germar, 1824)
Tribe DROMIINI
DROMIUS Bonelli, 1810
Subgenus MANODROMIUS Reitter, 1905
linearis (Olivier, 1795)
Subgenus PARADROMIUS Fowler, 1887
longiceps Dejean, 1826
Subgenus DROMIUS sensu stricto
agilis (Fabricius, 1787)
bimaculatus Dejean, 1825
angustus Brullé, 1834
meridionalis Dejean, 1825
discus Puel, 1919
quadrimaculatus (Linnaeus, 1758)
Subgenus CALODROMIUS Reitter, 1905
spilotus (Illiger, 1798)
quadrinotatus (Zenker in Panzer, 1800) non Fabricius, 1798
Subgenus PHILORHIZUS Hope, 1838
DROMIOLUS Reitter, 1905
melanocephalus Dejean, 1825
notatus Stephens, 1827
nigriventris Thomson, C.G., 1857
quadrisignatus Dejean, 1825
sigma (Rossi, 1790)
vectensis Rye, 1872
insignis sensu auctt. non Lucas, 1846
SYNTOMUS Hope, 1838
METABLETUS Schmidt-Göbel, 1846
foveatus (Fourcroy, 1785)
foveola (Gyllenhal, 1810)
obscuroguttatus (Duftschmid, 1812)
atratus (Dejean, 1825)
truncatellus (Linnaeus, 1761)
MICROLESTES Schmidt-Göbel, 1846
BLECHRUS Motschulsky, 1847
maurus (Sturm, 1827)
glabratus sensu auctt. non Duftschmid, 1812
minutulus (Goeze, 1777) 22
LIONYCHUS Wissmann, 1846
quadrillum (Duftschmid, 1812)
bipunctatus (Heer, 1838)
unicolor Schilsky, 1888
Tribe CYMINDINI
CYMINDIS Latreille, 1806
Subgenus CYMINDIS sensu stricto
angularis (Gyllenhal, 1810) †
axillaris (Fabricius, 1794)
macularis Fischer von Waldheim, 1824 23
Subgenus TARULUS Bedel, 1906
vaporariorum (Linnaeus, 1758)
basalis Gyllenhal, 1810
Tribe ZUPHIINI
POLISTICHUS Bonelli, 1810
connexus (Fourcroy, 1785)
fasciolatus sensu auctt. non Rossi, 1790
vittatus Brullé, 1834
Tribe DRYPTINI
DRYPTA Latreille, 1796
dentata (Rossi, 1790)
emarginata (Olivier, 1795)
1 Added by Luff (1990).
2 Temporary introduction, extinct; 19th-Century records only, Isles of Scilly.
3 Added by Luff (1989) and more fully by Anderson & Luff (1994).
4 Resident introduction, originating from N. Africa but now cosmopolitan; in sea-ports.
5 Forgotten name.
6 Temporary native; only 1 19th-Century record, then several in 20th Century from S.E. & N. England & Scotland, but only 1 record since 1970 (Berkshire).
7 Added by Anderson (1985).
8 Former resident native, found on banks of R. Clyde below Glasgow, Renfrewshire, to 1914.
9 Rare immigrant?; 2 specimens from London in 1950s.
10 Temporary native or introduction, extinct?; known only from Isle of Wight, last recorded from Sandown in 1914.
11 Temporary introduction; several 18th- and 19th-Century records from S. England, last recorded from Streatley (Berkshire) in 1905. Rediscovered near Whitehaven (Cumberland) in 1994, probably introduced with ships' ballast or timber.
12 Rare immigrant; several doubtful 19th-Century records, 1 confirmed record from Wimbledon (Surrey) in 1995 (Owen, 1996).
13 Rare immigrant.
14 Added by Telfer (2001b).
15 Rare immigrant or former resident native, extinct; several very old records from S. English coastal counties from Cornwall to E. Kent, last recorded from Isle of Wight c.1839.
16 Former resident native, extinct; coastal Dorset from Isle of Portland to St Alban's Head, last recorded in 1926, also an unconfirmed record from Hollingbury Hill (E. Sussex) in 1945. Also a very rare introduction (Kew Gardens, Surrey), but not certainly established.
17 Temporary introduction, from North America; 19th-Century records only from Gourock (Renfrewshire).
18 Former resident native, now probably extinct; known from Dorset, Isle of Wight and E. Sussex, last recorded from Charmouth (Dorset) in 1930.
19 Resident introduction, originating from S. Asia but now cosmopolitan.
20 Rare immigrant or former resident native, extinct; 2 19th-Century records, from near Devizes (Wiltshire) and Netley (Shropshire).
21 Former resident native, extinct; 18th/19th-Century records from Oakhampton Park (S. Somerset), Regent's Park (Middlesex)?, and Guestling (E. Sussex).
22 Added by Eversham & Collier (1997).
23 Added by Hammond (1982).
BRACHINUS Weber, 1801
Subgenus BRACHYNIDIUS Reitter, 1919
sclopeta (Fabricius, 1792) 1 Extinct?
Subgenus BRACHINUS sensu stricto
crepitans (Linnaeus, 1758) Bombadier Beetle
1 Former resident native, extinct?; several 19th-Century records from S. English coastal counties from Devon to Essex, last recorded Beachy Head (E. Sussex) in 1928.
Last updated: 28 February 2002
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